Gramma, San Diego Zoo’s ‘queen’ tortoise, dies at an estimated age of 141

Name: Gramma, San Diego Zoo’s ‘queen’ tortoise, dies at an estimated age of 141
Uploaded: 2025-11-25T17:09:09
Duration: PT29.0S
Description: Gramma, a Galápagos tortoise and the oldest resident of the San Diego Zoo, died Nov. 20. Her estimated 141-year lifespan spanned two world wars and 20 U.S. presidents.

Gramma, a Galápagos tortoise and the oldest resident of the San Diego Zoo, died Nov. 20. Her estimated 141-year lifespan spanned two world wars and 20 U.S. presidents.

Gramma, San Diego Zoo’s ‘queen’ tortoise, dies at an estimated age of 141

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